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Bat

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For other uses, see Bat=20 (disambiguation).
Bat
Temporal range: = 52=E2=80=930 Ma
Pre=D0=84
=D0=84
O
S
D
C
P
T
J
K
Pg
N
Early Eocene=20 =E2=80=93 Recent
Townsend= 's=20 big-eared bat, Corynorhinus townsendii
Scientifi= c=20 classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Superorder: Laurasiatheria[1]
Order: Chiroptera
Blumenb= ach,=20 1779
Suborders

See article

Worldwide=20 distribution of bat species

Bats are mammals of the order=20 Chiroptera (3D"play"=20 /ka=C9=AA=CB=88r=C9=92pt=C9=99r=C9=99/;=20 from the Greek=20 =CF=87=CE=B5=CE=AF=CF=81 - cheir, "hand"[2]=20 and =CF=80=CF=84=CE=B5=CF=81=CF=8C=CE=BD - pteron, "wing"[3])=20 whose forelimbs form webbed wings,=20 making them the only mammals naturally capable of true and sustained flight. By contrast, = other=20 mammals said to fly, such as flying = squirrels, gliding=20 possums, and colugos, glide rather = than fly,=20 and can only glide for short distances. Bats do not flap their entire = forelimbs,=20 as birds do, but=20 instead flap their spread-out digits,[4]=20 which are very long and covered with a thin membrane=20 or patagium.

Bats represent about 20% of all classified mammal species worldwide, = with=20 about 1,240 bat species divided into two suborders: the less specialized = and=20 largely fruit-eating 'megachiroptera', or flying foxes, and the more = highly=20 specialized and echolocating 'microchiroptera'.[5]=20 About 70% of bats are insectivores.=20 Most of the rest are frugivores,=20 or fruit eaters. A few species, such as the fish-eating = bat, feed=20 from animals other than insects, with the vampire=20 bats being the only parasitic=20 mammalian species. Bats are present throughout most of the world, = performing=20 vital ecological roles of pollinating=20 flowers and=20 dispersing fruit seeds. Many tropical plant species depend entirely on = bats for=20 the distribution of their seeds. Bats are important in eating insect = pests,=20 reducing the need for pesticides. The smallest bat is the Kitti's=20 hog-nosed bat, measuring 29=E2=80=9334 mm (1.14=E2=80=931.34 in) in = length, 15 cm=20 (5.91 in) across the wings and 2=E2=80=932.6 g (0.07=E2=80=930.09 = oz) in mass.[= 6][7]=20 It is also arguably the smallest extant species of mammal, with the Etruscan shrew = being the=20 other contender.[= 8]=20 The largest species of bat is the gia= nt=20 golden-crowned flying fox, which is 336=E2=80=93343 mm = (13.23=E2=80=9313.50 in) long, has a=20 wingspan of 1.6 m (5 ft 3 in) and weighs = approximately=20 1.1=E2=80=931.2 kg (2=E2=80=933 lb).[9]

Contents

[edit]=20 Classification and=20 evolution

<= IMG=20 class=3D"thumbimage" alt=3D"" = src=3D"http://upload.wikimedia.org/wikipedia/commons/3/3f/Golden_crowned_= fruit_bat.jpg"=20 width=3D"200" height=3D"319">
<= IMG=20 alt=3D"" = src=3D"http://bits.wikimedia.org/static-1.20wmf8/skins/common/images/magn= ify-clip.png"=20 width=3D"15" height=3D"11">
Gia= nt=20 golden-crowned flying fox, Acerodon jubatus

Bats are mammals. They are often = mistakenly called "flying rodents"=20 or "flying rats". In many languages, the word for "bat" is cognate with = the word=20 for "mouse": for example, chauve-souris ("bald-mouse") in French, = murci=C3=A9lago ("blind mouse") in Spanish, = =D0=BB=D0=B5=D1=82=D1=83=D1=87=D0=B0=D1=8F =D0=BC=D1=8B=D1=88=D1=8C = ("flying mouse") in=20 Russian, and nahkhiir ("leather mouse") in Estonian, = vlermuis=20 (winged mouse) in Afrikaans, from the Dutch word vleermuis. An = older=20 English name for bats is flittermice,=20 which matches their name in other Germanic=20 languages (for example German=20 Fledermaus and Swedish=20 fladdermus).[10]=20 However, they are not directly related to rodents, and much less to = birds, and=20 do not in fact have any closely related orders (their uniqueness can be=20 demonstrated by the fact their closest living genetic relatives are = thought to=20 be carnivorans, certain = hoofed animals, such = as alpacas and hippopotamuses, = and sea=20 mammals, such as dolphins.)

The two traditionally recognized suborders of bats are:

Not all megabats are larger than microbats. The major distinctions = between=20 the two suborders are:

Megabats eat fruit, nectar=20 or pollen,=20 while most microbats eat insects;=20 others may feed on the blood=20 of animals, small mammals, fish, frogs, fruit, pollen or nectar. = Megabats have=20 well-developed visual=20 cortices and show good visual=20 acuity, while microbats rely on echolocation=20 for navigation and finding prey.

The phylogenetic relationships of the different groups of bats have = been the=20 subject of much debate. The traditional subdivision between = Megachiroptera and=20 Microchiroptera reflects the view that these groups of bats have evolved = independently of each other for a long time, from a common ancestor = already=20 capable of flight. This hypothesis recognized differences between = microbats and=20 megabats and acknowledged that flight has only evolved once in mammals. = Most=20 molecular biological evidence supports the view that bats form a single = or=20 monophyletic group.= [11]

Researchers have proposed alternate views of chiropteran phylogeny = and classific= ation,=20 but more research is needed.

Genetic=20 evidence indicates megabats originated during the early Eocene = and should be placed within the four major lines of microbats.

Consequently, two new suborders based on molecular data have been = proposed.=20 The new suborder Yinpterochiropte= ra=20 includes the Pteropodidae or = megabat family,=20 as well as the Rhinolophidae,=20 Hipposideridae, = Craseonycteridae, Megadermatidae, = and Rhinopomatidae = families.=20 The new suborder Yangochiroptera= =20 includes all the remaining families of bats (all of which use laryngeal=20 echolocation). These two new suborders are strongly supported by = statistical=20 tests. Teeling (2005) found 100% bootstrap support in all maximum = likelihood=20 analyses for the division of Chiroptera into these two modified = suborders. This=20 conclusion is further supported by a 15-base-pair deletion in BRCA1 and = a=20 seven-base-pair deletion in PLCB4 present in all Yangochiroptera and = absent in=20 all Yinpterochiroptera.[12]=20 The chiropteran phylogeny based on molecular evidence is controversial = because=20 microbat paraphyly implies one of two seemingly unlikely hypotheses = occurred.=20 The first suggests laryngeal echolocation evolved twice in Chiroptera, = once in=20 Yangochiroptera and once in the rhinolophoids.[13][14]=20 The second proposes laryngeal echolocation had a single origin in = Chiroptera,=20 was subsequently lost in the family Pteropodidae (all megabats), and = later=20 evolved as a system of tongue-clicking in the genus = Rousettus.[15]

Common = pipistrelle,=20 Pipistrellus pipistrellus

Analyses of the sequence of the "vocalization" gene, FoxP2 was = inconclusive as to whether laryngeal echolocation was secondarily lost = in the=20 pteropodids or independently gained in the echolocating lineages.[16]=20 However, analyses of the "hearing" gene, Prestin seemed to favor = the=20 independent gain in echolocating species rather than a secondary loss in = the=20 pteropodids.[17]

In addition to Yinpterochiroptera and Yangochiroptera, the names=20 Pteropodiformes and Vespertilioniformes have also been proposed for = these=20 suborders.[18][19]=20 Under this new proposed nomenclature, the suborder Pteropodiformes = includes all=20 extant bat families more closely related to the genus Pteropus = than the=20 genus Vespertilio, while the suborder Vespertilioniformes = includes all=20 extant bat families more closely related to the genus Vespertilio = than to=20 the genus Pteropus.

In the 1980s, a hypothesis based on morphological<= /A>=20 evidence was offered that stated the Megachiroptera evolved flight = separately=20 from the Microchiroptera. The so-called flying=20 primates theory proposed when adaptations to flight are removed, the = Megachiroptera are allied to primates=20 by anatomical features not shared with Microchiroptera. One example is = the=20 brains of megabats show a number of advanced characteristics that link = them to=20 primates. Although recent genetic studies support the monophyly of = bats,[20]=20 debate continues as to the meaning of available genetic and = morphological=20 evidence.[21]

Little fossil evidence is available to help map the evolution of bats, = since their=20 small, delicate skeletons do not = fossilize very=20 well. However, a Late=20 Cretaceous tooth from South America resembles that of an early=20 microchiropteran bat. The oldest known, definitely identified bat = fossils, such=20 as Icaronycteris= , Archaeonycteris= ,=20 Palaeochiropteryx= =20 and Hassianycteris,=20 are from the early Eocene period, 52.5 million years ago.= [11]=20 These fossil bats were already very similar to modern microbats.=20 Archaeopteropus, formerly classified as the earliest known=20 megachiropteran, is now classified as a microchiropteran.

Bats were formerly grouped in the superorder Archonta=20 along with the treeshrews=20 (Scandentia), colugos (Dermoptera), = and the primates,=20 because of the apparent similarities between Megachiroptera and such = mammals.=20 Genetic studies have now placed bats in the superorder Laurasiatheria = along with=20 carnivorans, pangolins, odd-toed = ungulates, even-toed=20 ungulates, and cetaceans.[1]


   Laurasiatheria&n= bsp;  

 Eulipotyphla


   Scrotifera &nbs= p; 

 Chiroptera

   Fereuungulata&nbs= p;  
   Ferae   

 Pholidota

 Carnivora

 Perissodactyla    

 Cetartiodactyla=





"Chiroptera" from Ernst Haeckel's = Kunstformen=20 der Natur, 1904

The traditional classification of bats is:

Megabats primarily eat fruit or nectar. In New Guinea, they are = likely to=20 have evolved for some time in the absence of microbats. This has = resulted in=20 some smaller megabats of the genus Nyctimene= =20 becoming (partly) insectivorous to fill the vacant microbat ecological = niche.=20 Furthermore, there is some evidence that the fruit bat genus Pteralopex from = the Solomon = Islands, and its=20 close relative Mirimiri=20 from Fiji, have=20 evolved to fill some niches that were open because there are no = nonvolant or=20 non-flying mammals in those islands.

[edit]=20 Fossil bats

There are few fossilized remains of bats, as they are terrestrial and = light-boned. An estimated only 12% of the bat fossil record is complete = at the=20 genus level.[22]=20 An Eocene bat,=20 Onychonycteris=20 finneyi, was found in the 52-million-year-old Green River=20 Formation in Wyoming, United States, in 2003.[23][24]=20 It had characteristics indicating it could fly,= yet the=20 well-preserved skeleton showed the cochlea=20 of the inner ear lacked development needed to support the greater = hearing=20 abilities of modern bats. This provided evidence flight in bats = developed well=20 before echolocation. The team that found the remains of this species, = named=20 Onychonycteris finneyi, recognized it lacked ear and throat = features=20 present not only in echolocating bats today, but also in other known prehistoric = species. Fossil=20 remains of another Eocene bat, Icaronycteris= ,=20 were found in 1960.

The appearance and flight movement of bats 52.5 million years ago = were=20 different from those of bats today. Onychonycteris had claws on = all five=20 of its fingers, whereas modern bats have at most two claws appearing on = two=20 digits of each hand. It also had longer hind legs and shorter forearms, = similar=20 to climbing mammals that hang under branches such as sloths=20 and gibbons.=20 This palm-sized bat had broad, short wings, suggesting it could not fly = as fast=20 or as far as later bat species. Instead of flapping its wings = continuously while=20 flying, Onychonycteris likely alternated between flaps and glides = while=20 in the air. Such physical characteristics suggest this bat did not fly = as much=20 as modern bats do, rather flying from tree to tree and spending most of = its=20 waking day climbing or hanging on the branches of trees.[25]

[edit]=20 Habitats

Flight has enabled bats to become one of the most widely distributed = groups=20 of mammals.[26]=20 Apart from the Arctic, the Antarctic and a few isolated oceanic islands, = bats=20 exist all over the world.[27]=20 Bats are found in almost every habitat available on Earth. Different = species=20 select different habitats during different seasons, ranging from = seasides to=20 mountains and even deserts, but bat habitats have two basic = requirements:=20 roosts, where they spend the day or hibernate, and places for foraging. = Bat=20 roosts can be found in hollows, crevices, foliage, and even human-made=20 structures, and include "tents" the bats construct by biting leaves.[28]

The United States is home to an estimated 45 to 48 species of = bats.[29][30]=20 The three most common species are Myotis = lucifugus=20 (little brown bat), Eptesicus = fuscus (big brown bat), and Tadarida=20 brasiliensis (Mexican free-tailed bat). The little and the big = brown bat=20 are common throughout the northern two-thirds of the country, while the = Mexican=20 free-tailed bat is the most common species in the southwest.[31]

[edit]=20 Anatomy

Skeleton of a Greater = mouse-eared=20 bat (Myotis myotis)
<= IMG=20 class=3D"thumbimage" alt=3D"" = src=3D"http://upload.wikimedia.org/wikipedia/commons/thumb/5/52/MyotisSca= pulaeRibs120810.jpg/175px-MyotisScapulaeRibs120810.jpg"=20 width=3D"175" height=3D"138">
<= IMG=20 alt=3D"" = src=3D"http://bits.wikimedia.org/static-1.20wmf8/skins/common/images/magn= ify-clip.png"=20 width=3D"15" height=3D"11">
Scapulae, spine and ribs of = Myotis=20 lucifugus (little brown bat)

[edit]=20 Echolocation

Main article: Animal=20 echolocation#Bats
Spectrogram=20 of Pipistrellus pipistrellus (Common=20 Pipistrelle) bat vocalizations - one echolocation=20 call, followed by series of multiharmonic pulses, forming a social call = (here,=20 precisely, an advertisement call, produced by territorial male during = the mating=20 season in autumn).[32]=20 This spectrogram was generated with Fatpigdog's PC-based real-time FFT=20 spectrum analyzer.
3D""
Pipistrellus = Pulses
Recording of=20 Pipistrellus pipistrellus bat time-expanded echolocation = calls and=20 social call.
Problems=20 listening to this file? See media=20 help.

Bat echolocation is a perceptual system where ultrasonic sounds are = emitted=20 specifically to produce echoes. By comparing the outgoing pulse with the = returning echoes, the brain and auditory nervous system can produce = detailed=20 images of the bat's surroundings. This allows bats to detect, localize = and even=20 classify their prey in complete darkness. At 130 decibels in intensity, = bat=20 calls are some of the most intense, airborne animal sounds.[33]

To clearly distinguish returning information, bats must be able to = separate=20 their calls from the echoes they receive. Microbats use two distinct=20 approaches.

  1. Low duty cycle echolocation: Bats can separate their calls and = returning=20 echos by time. Bats that use this approach time their short calls to = finish=20 before echoes return. This is important because these bats contract = their=20 middle ear muscles when emitting a call so they can avoid deafening=20 themselves. The time interval between call and echo allows them to = relax these=20 muscles so they can clearly hear the returning echo.[34]=20 The delay of the returning echos provides the bat with the ability to = estimate=20 range to their prey.
  2. High duty cycle echolocation: Bats emit a continuous call and = separate=20 pulse and echo in frequency. The ears of these bats are sharply tuned = to a=20 specific frequency range. They emit calls outside of this range to = avoid=20 self-deafening. They then receive echoes back at the finely tuned = frequency=20 range by taking advantage of the Doppler shift = of their=20 motion in flight. The Doppler shift of the returning echos yields = information=20 relating to the motion and location of the bat's prey. These bats must = deal=20 with changes in the Doppler shift due to changes in their flight = speed. They=20 have adapted to change their pulse emission frequency in relation to = their=20 flight speed so echoes still return in the optimal hearing range.[35]

The new Yinpterochiroptera and Yangochiroptera classification of = bats,=20 supported by molecular evidence, suggests two possibilities for the = evolution of=20 echolocation. It may have been gained once in a common ancestor of all = bats and=20 was then subsequently lost in the Old World fruit bats, only to be = regained in=20 the horseshoe bats, or echolocation evolved independently in both the=20 Yinpterochiroptera and Yangochirpotera lineages.[36]

Two groups of moths exploit a bat sense = to=20 echolocate: tiger moths produce = ultrasonic signals = to warn=20 the bats they (the moths) are chemically protected or aposematic. This = was once=20 thought to be the biological equivalent of "radar jamming", = but this=20 theory has yet to be confirmed. The moths Noctuidae=20 have a hearing organ called a tympanum,=20 which responds to an incoming bat signal by causing the moth's flight = muscles to=20 twitch erratically, sending the moth into random evasive maneuvers.

In addition to echolocating prey, bat ears are sensitive to the = fluttering of=20 moth wings, the sounds produced by tymbalate=20 insects and the movement of ground-dwelling prey, such as centipedes, = earwigs,=20 etc. The complex geometry of ridges on the inner surface of bat ears = helps to=20 sharply focus not only echolocation signals, but also to passively = listen for=20 any other sound produced by the prey. These ridges can be regarded as = the=20 acoustic equivalent of a fresnel=20 lens, and may be seen in a large variety of unrelated animals, such = as the=20 aye-aye, lesser=20 galago, bat-eared=20 fox, mouse lemur and = others.[37][38][39]

By repeated scanning, bats can mentally construct an accurate image = of the=20 environment in which they are moving or their prey item.[40]

[edit]=20 Other senses

Although the eyes=20 of most microbat species are small and poorly developed, leading to poor = visual=20 acuity, none of them are blind. Microbats use vision to navigate, = especially=20 for long distances when beyond the range of echolocation. Some species = have been=20 shown to be able to detect ultraviolet=20 light. They also have high-quality senses of smell and hearing. Bats = hunt at=20 night to avoid competition with birds, and travel large distances of up = to=20 800 km in their search for food.[4]

[edit]=20 Wings

A preserved fruit bat showing how = the skeleton=20 fits inside its skin.
Thermographic image of a bat using = trapped air=20 as insulation

The finger bones of bats are much more flexible than those of other = mammals,=20 owing to their flattened cross-section and to low levels of minerals = such as=20 calcium near their tips. The skin on their wing membranes has more = elasticity,=20 so can stretch much more than other mammals.

The wings of bats are much thinner than those of birds, allowing bats = to=20 maneuver more quickly and more accurately than birds.[citation=20 needed] It is also delicate, ripping easily;[41]=20 however, the tissue of the bat's membrane is able to regrow, such that = small=20 tears can heal quickly.[41][42]=20 The surface of their wings is equipped with touch-sensitive receptors on = small=20 bumps called Merkel=20 cells, also found on human fingertips. These sensitive areas are = different=20 in bats, as each bump has a tiny hair in the center,= [43]=20 making it even more sensitive and allowing the bat to detect and collect = information about the air flowing over its wings, and to fly more = efficiently by=20 changing the shape of its wings in response.= [43]=20 An additional kind of receptor cell is found in the wing membrane of = species=20 that use their wings to catch prey. This receptor cell is sensitive to = the=20 stretching of the membrane.= [43]=20 The cells are concentrated in areas of the membrane where insects hit = the wings=20 when the bats capture them.

[edit]=20 Other

The teeth of microbats resemble insectivorans.=20 They are very sharp to bite through the hardened armor = of insects=20 or the skin of fruit.

Mammals have one-way valves in veins=20 to prevent the blood from flowing backwards, but bats also have one-way = valves=20 in arteries.

The tube-lipped = nectar bat (Anoura fistulata), has the longest tongue of any mammal = relative to=20 its body size. This is beneficial to them in terms of pollination and = feeding.=20 Its long, narrow tongues can reach deep into the long cup shape of some = flowers. When the=20 tongue retracts, it coils up inside its rib cage.[44]

[edit]=20 Behaviour

Most microbats are nocturnal[45]=20 and are active at twilight. A large portion of bats migrate = hundreds of=20 kilometres to winter hibernation dens,[= 46]=20 some pass into torpor in cold weather, = rousing=20 and feeding when warm weather allows for insects to be active.[47]=20 Others retreat to caves for winter and hibernate=20 for six months.[47]=20 Bats rarely fly in rain, as the rain interferes with their echo = location, and=20 they are unable to locate their food.

The social structure of bats varies, with some bats leading solitary = lives=20 and others living in caves colonized by more than a million bats.[48]=20 The fission-fusi= on=20 social structure is seen among several species of bats. The term = "fusion"=20 refers to a large numbers of bats that congregate in one roosting area, = and=20 "fission" refers to breaking up and the mixing of subgroups, with = individual=20 bats switching roosts with others and often ending up in different trees = and=20 with different roostmates.

Studies also show bats make all kinds of sounds to communicate with = others.=20 Scientists in the field have listened to bats and have been able to = identify=20 some sounds with some behaviour bats will make after the sounds are = made.[48]

Insectivores make up 70% of bat species and locate their prey by = means of echolocati= on.=20 Of the remainder, most feed on fruits.[49]=20 Only three species sustain themselves with blood.=20 Some species even prey on vertebrates:=20 these are the leaf-nosed=20 bats (Phyllostomidae) of Central=20 America and South=20 America, and the two bulldog=20 bat (Noctilionidae) species, which feed on fish.=20 At least two species of bat are known to feed on bats: the spectral bat, = also known as=20 the American false vampire bat, and the ghost=20 bat of Australia.[49]=20 One species, the greater=20 noctule bat, catches and eats small birds=20 in the air.

Predators of bats include bat=20 hawks and bat=20 falcons.

[edit]=20 Reproduction

Newborn common pipistrelle, = Pipistrellus=20 pipistrellus
Colony of mouse-eared = bats,=20 Myotis myotis

Most bats have a breeding season, which is in the spring for species = living=20 in a temperate climate. Bats may have one to three litters in a season,=20 depending on the species and on environmental conditions, such as the=20 availability of food and roost sites. Females generally have one = offspring at a=20 time, which could be a result of the mother's need to fly to feed while=20 pregnant. Female bats nurse their young until they are nearly adult = size,=20 because a young bat cannot forage on its own until its wings are fully=20 developed.

Female bats use a variety of strategies to control the timing of = pregnancy=20 and the birth of young, to make delivery coincide with maximum food = ability and=20 other ecological factors. Females of some species have delayed = fertilization, in=20 which sperm are stored in the reproductive tract for several months = after=20 mating. In many such cases, mating occurs in the fall, and fertilization = does=20 not occur until the following spring. Other species exhibit delayed=20 implantation, in which the egg is fertilized after mating, but remains = free in=20 the reproductive tract until external conditions become favorable for = giving=20 birth and caring for the offspring.

In yet another strategy, fertilization and implantation both occur, = but=20 development of the fetus is delayed until favorable conditions prevail, = during=20 the delayed development the mother still gives the fertilized egg = nutrients, and=20 oxygenated blood to keep it alive. However this process can go for a = long period=20 of time, because of the advanced gas exchange system[50].=20 All of these adaptations result in the pup being born during a time of = high=20 local production of fruit or insects.

At birth, the wings are too small to be used for flight. Young microbats become = independent at=20 the age of six to eight weeks, while megabats=20 do not until they are four months old.

A single bat can live over 20 years, but the bat population growth is = limited=20 by the slow birth rate.[51]

[edit]=20 Hunting,=20 feeding, and drinking

<= IMG=20 alt=3D"" = src=3D"http://bits.wikimedia.org/static-1.20wmf8/skins/common/images/magn= ify-clip.png"=20 width=3D"15" height=3D"11">
A very young bat in Tamil=20 Nadu

Newborn bats rely on the milk from their mother.[52]=20 When they are a few weeks old, bats are expected to fly and hunt on = their own.=20 It is up to them to find and catch their prey, along with satisfying = their=20 thirst.[53]

[edit]=20 Hunting

Most bats are nocturnal creatures. Their daylight hours are spent = grooming=20 and sleeping; they hunt during the nighttime hours. The means by which = bats=20 navigate while finding and catching their prey in the dark was unknown = until the=20 1790s, when Lazzaro=20 Spallanzani conducted a series of experiments on a group of blind = bats.=20 These bats were placed in a room in total darkness, with silk threads = strung=20 across the room. Even then, the bats were able to navigate their way = through the=20 room. Spallanzani concluded the bats were not using their eyes to fly = through=20 complete darkness, but something else.

Spallanzani decided the bats were able to catch and find their prey = through=20 the use of their ears. To prove this theory, Spallanzani plugged the = ears of the=20 bats in his experiment. To his pleasure, he found the bats with plugged = ears=20 were not able to fly with the same amount of skill and precision they = were able=20 to without their ears plugged.

Bats seem to use their ears to locate and catch their prey, but how = they=20 accomplish this was not discovered until the 1930s, by Donald R. = Griffin, a=20 biology student at Harvard College at the time. He discovered bats use=20 echolocation to locate and catch their prey. When bats fly, they produce = a=20 constant stream of high-pitched sounds only bats are able to hear. When = the=20 sound waves produced by these sounds hit an insect or other animal, the = echoes=20 bounce back to the bat, and guide them to the source.[53]

[edit]=20 Feeding and = diet

The majority of food consumed by bats includes insects, fruits and = flower=20 nectar, vertebrates and blood.[= 54]=20 Almost three-fourths of the world=E2=80=99s bats are insect eaters. = Insects consumed by=20 bats include both aerial and ground-dwelling insects. Each bat is = typically able=20 to consume one-third of its body weight in insects each night, and = several=20 hundred insects in a few hours. This means a group of one thousand bats = could=20 eat four tons of insects each year. If bats were to become extinct, the = insect=20 population is calculated to reach an alarmingly high number.[= 55]

[edit]=20 Vitamin C

In a test of 34 bat species from six major families of bats, = including major=20 insect- and fruit-eating bat families, bats in all tested families were = found to=20 have lost the ability to make vitamin C, and this loss may derive from a = common=20 bat ancestor, as a single mutation.[56]

[edit]=20 Aerial=20 insectivores

Watching a bat catch and eat an insect is difficult. The action is so = fast=20 that all one sees is a bat rapidly change directions, and continue on = its way.=20 Scientist Frederick A. Webster discovered how bats catch their prey. In = 1960,=20 Webster developed a high-speed camera that was able to take one thousand = pictures per second. These photos revealed the fast and precise way in = which=20 bats catch insects.[53]=20 Occasionally, a bat will catch an insect in mid-air with its mouth, and = eat it=20 in the air. However, more often than not, a bat will use its tail = membrane or=20 wings to scoop up the insect and trap it in a sort of "bug net".[52]=20 Then, the bat will take the insect back to its roost. There, the bat = will=20 proceed to eat said insect, often using its tail membrane as a kind of = napkin,=20 to prevent its meal from falling to the ground.[= 54]

[edit]=20 Forage = gleaners

These bats typically fly down and grasp their prey off the ground = with their=20 teeth, and take it to a nearby perch to eat it. Generally, these bats do = not use=20 echolocation to locate their prey. Instead, they rely on the sounds = produced by=20 the insects. Some make unique sounds, and almost all make some noise = while=20 moving through the environment.[52]

[edit]=20 Fruits and = flower=20 nectar
A colony of great = fruit-eating=20 bats

Fruit eating, or frugivory, is a specific habit found in two families = of=20 bats. Megachiropterans and microchiropterans both include species of bat = that=20 feed on fruits. These bats feed on the juices of sweet fruits, and = fulfill the=20 needs of some seeds to be dispersed. The fruits preferred by most = fruit-eating=20 bats are fleshy and sweet, but not particularly strong smelling or = colorful.[52]=20 To get the juice of these fruits, bats pull the fruit off the trees with = their=20 teeth, and fly back to their roosts with the fruit in their mouths. = There, the=20 bats will consume the fruit in a specific way. To do this, the bats = crush open=20 the fruit and eat the parts that satisfy their hunger. The remainder of = the=20 fruit; the seeds and pulp, are spat onto the ground. These seeds take = root and=20 begin to grow into new fruit trees.[= 55]=20 =E2=80=9COver one hundred and fifty types of plants depend on bats in = order to=20 reproduce=E2=80=9D.[= 55]

Some bats prefer the nectar of flowers to insects or other animals. = These=20 bats have evolved specifically for this purpose. For example, these bats = possess=20 long muzzles and long, extensible tongues covered in fine bristles that = aid them=20 in feeding on particular flowers and plants.[52]=20 When they sip the nectar from these flowers, pollen gets stuck to their = fur, and=20 is dusted off when the bats take flight, thus pollinating the plants = below=20 them.[= 55]=20 The rainforest is said to be the most benefitted of all the biomes where = bats=20 live, because of the large variety of appealing plants.[57]=20 Because of their specific eating habits, nectar-feeding bats are more = prone to=20 extinction than any other type of bat.[58]=20 However, bats benefit from eating fruits and nectar just as much from = eating=20 insects.[59]

[edit]=20 Vertebrates

A small group of carnivorous bats feed on other vertebrates and are=20 considered the top carnivores of the bat world.[52]=20 These bats typically eat a variety of animals, but normally consume = frogs,=20 lizards, birds, and sometimes other bats.[= 55]=20 For example, one vertebrate predator, Trachops cirrhosus, is = particularly=20 skilled at catching frogs. These bats locate large groups of frogs by=20 distinguishing their mating calls from other sounds around them. They = follow the=20 sounds to the source and pluck them from the surface of the water with = their=20 sharp canine teeth.[52]=20 Another example is the greater=20 noctule bat, which is believed to catch birds on the wing.

Also, several species of bat feed on fish. These types of bats are = found on=20 almost all continents. They use echolocation to detect tiny ripples in = the=20 water=E2=80=99s surface to locate fish. From there, the bats swoop down = low, inches from=20 the water, and use specially enlarged claws on their hind feet to grab = the fish=20 out of the water. The bats then take the fish to a feeding roost and = consume the=20 animal.[52]

[edit]=20 Blood

A few species of bats consume blood exclusively as their diet. This = type of=20 diet is referred to as hematophagy, and three species of bats exhibit = this=20 behavior. These species are the common,=20 the white-wing= ed,=20 and the hairy-legg= ed=20 vampire bats. The common vampire bat typically consumes the blood of = mammals,=20 while the hairy-legged and white-winged vampires feed on the blood of = birds.[60]

[edit]=20 Results of = eating

Bats' dung, or guano, is so rich in = nutrients=20 that it is mined from caves, bagged, and used by farmers to fertilize = their=20 crops. During the U.S.=20 Civil War, guano was used to make gunpowder.[= 55]

To survive hibernation=20 months, some species build up large reserves of body fat, both as fuel = and as=20 insulation.[53]

[edit]=20 Drinking

Generally, bats drink water. In 1960, Frederic A. Webster discovered = how bats=20 are able to acquire this water. To do this, Webster developed a = high-speed=20 camera and flashgun that could take 1000 photos per second. Webster's = camera=20 captured the bat's method of skimming the surface of a body of water, = and=20 lowering its jaw to get just one drop of water. It then skims again to = get a=20 second drop of water, and so on, until it has had its fill of water. Its = precision and control is very fine, and it almost never misses.[53]=20 Other, such as the flying=20 fox or fruit=20 bat, gently skim the water's surface, then land nearby to lick water = from=20 their chest fur.[61]

[edit]=20 Conservation=20 efforts

Through conservancy efforts of groups such as the O= rganization=20 for Bat Conservation and Bat Conservation International, bats are = becoming=20 better understood and people beginning to understand the crucial role = bats play=20 in insect control and pollination.

In the United=20 Kingdom, all bats are protected under the Wildli= fe and=20 Countryside Acts, and even disturbing a bat or its roost can be = punished=20 with a heavy fine.

In Sarawak, Malaysia, bats are = protected=20 species under the Wildlife Protection Ordinance 1998 (see Malaysian = Wildlife=20 Law). The large naked bat (see Mammals=20 of Borneo) and greater nectar bat are consumed by the local = communities.

Bats can be a tourist attraction. The Congress=20 Avenue Bridge in Austin,=20 Texas = is the=20 summer home to North=20 America's largest urban bat colony, an estimated 1,500,000 Mexican=20 free-tailed bats, which eat an estimated 10,000 to 30,000 pounds of = insects=20 each night. An estimated 100,000 tourists per year visit the bridge at = twilight=20 to watch the bats leave the roost.

[edit]=20 Artificial = roosts

Very large bat house, Tallahassee, = Florida,=20 United States

Many people put up bat houses to attract bats just as some put up birdhouses to attract = birds.=20 Reasons for this vary, but most center around the bats being the primary = nocturnal insectivores in most, if not all, ecologies. Bat houses can be = made=20 from scratch or from kits, or bought ready made. Plans for bat houses = exist on=20 many web sites, as well as guidelines for designing a bat house.[62]=20 Some conservation societies give away free bat houses to enthusiasts=20 worldwide.[citation=20 needed]

Constructed in March of 1991, the University=20 of Florida bat house is the largest occupied artificial roost in the = world.=20 The bat house has an approximated 300,000 insect-eating residents. These = bats=20 can eat upwards of 2.5 billion bugs(2,500 pounds) per night.

An unfortunate fire in 1987 caused the colony of bats (residing in = Johnson=20 Hall's attic) to become homeless. This forced them to move to the James = G.=20 Pressly Stadium on the north side of campus. The odor and the guano from = this=20 newly arrived colony did not please spectators, thus furthering the = movement for=20 a new bat-ordained structure. In September of 1991, thousands of bats = were=20 captured and transported to the newly built bat house. In the following=20 evenings, these bats flew away, and found new homes. For three years the = bat=20 house remained empty. Finally, in 1995, the bats moved in permanently, = and the=20 colony continued to grow. The colony grew so much, that in 2009, part of = the=20 original bat house collapsed, and a new =E2=80=9CBat Barn=E2=80=9D was = constructed next to it.=20 Along with that, the interior of the original was rebuilt. [63]

In Britain, British=20 hardened field defences of World War II have been converted to make = roosts=20 for bats. Pillboxes that are well dug-in and thick-walled are naturally = damp and=20 provide the stable thermal environment required by bats that would = otherwise=20 hibernate in caves. With a few minor modifications, suitable pillboxes = can be=20 converted to artificial caves for bats.[64][65]

Again in the UK, purpose-built bat houses are occasionally built when = existing roosts are destroyed by developments such as new roads; one = such has=20 been built associated with bat=20 bridges on the new (2008) A38 Dobwalls=20 bypass.[citation=20 needed]

[edit]=20 Threats

A little=20 brown bat with white nose syndrome

While conservation efforts are in place to protect bats, many threats = still=20 remain.

[edit]=20 White nose=20 syndrome

Main article: White nose=20 syndrome

White nose syndrome is a condition associated with the deaths of more = than a=20 million bats in the northeas= tern=20 United States.= [66]=20 The disease is named after a white fungus,=20 Geomyces destructans, found growing on the muzzles, ears, and wings of = afflicted=20 bats. This fungus, which can be spread from bat to bat, is the sole = cause of the=20 disease.[67]=20 Mortality rates of 90=E2=80=93100% have been observed in some caves.[68]=20 At least six species of hibernating bats are affected, including the = endangered=20 Indiana=20 bat.[67]=20 Because the affected species have long lifespans, and birth=20 rates of only about one offspring per year, populations are not = expected to=20 recover quickly.[67]

[edit]=20 Barotrauma and wind=20 turbines

Although sounds or vibrations emitted from wind turbines have no = adverse=20 effect on human health,[69]=20 evidence suggests barotrauma=20 is causing bat fatalities around wind farms.[70]=20 The lungs of bats are typical mammalian=20 lungs, and unlike the=20 lungs of birds, they are thought to be more sensitive to sudden air = pressure=20 changes in their immediate vicinity, such as near wind=20 turbines, and are more liable to rupture.[= 71]=20 Bats suffer a higher death rate than birds in the neighborhood of wind = turbines.[72][73][74]=20 Since there are no signs of external trauma, the cause has been = hypothesized to=20 be a greater sensitivity to sudden pressure fluctuations in the = mammalian lung=20 than in that of birds.[75]=20 In addition, it has been suggested that bats are attracted to these = structures,=20 perhaps seeking roosts, and thereby increasing the death rate.[= 71]

[edit]=20 Pathogens=20 and role in the transmission of zoonoses

Among ectoparasites,=20 bats occasionally carry fleas, but are one of the = few=20 mammalian orders that cannot host lice=20 (most of the others are water animals).

Bats are natural=20 reservoirs for a large number of zoonotic=20 pathogens,[76]=20 including rabies,<= SUP=20 id=3D"cite_ref-McColl2000_76-0" class=3D"reference">[= 77]=20 s= evere=20 acute respiratory syndrome (SARS),[78]=20 Henipavirus (i.e. = Nipah=20 virus and Hendra virus)[= 79]=20 and possibly ebola=20 virus.[<= /SPAN>80][<= /SPAN>81]=20 Their high mobility, broad distribution, and social behaviour (communal = roosting=20 and fission-fusion social structure) make bats favourable hosts and = vectors of=20 disease. Many species also appear to have a high tolerance for = harbouring=20 pathogens and often do not develop disease while infected. However, = contrary to=20 folklore, this is not true of rabies, which is as fatal to bats as it is = to all=20 other species. However, a bat may be ill with rabies for a longer time = than=20 other mammals.[citation=20 needed]

In regions where rabies is endemic, only 0.5% of bats carry the = disease. In=20 the United States, = bats=20 typically constitute around a quarter of reported cases of rabies in = wild=20 animals. However, their bites=20 account for the vast majority of cases of rabies=20 in humans. Of the 36 cases of domestically-acquired rabies recorded in = the=20 country in 1995-2010, two were caused by dog bites and four patients = were=20 infected by receiving transplants from an organ donor who had previously = died of=20 rabies. All other cases were caused by bat bites.[82]=20 Rabies is considered fully preventable if the patient is administered a = vaccine=20 prior to the onset of symptoms. However, unlike raccoon=20 or skunk bites,=20 bat bites may go ignored or unnoticed and hence untreated. Rabid bats = are=20 broadly distributed throughout the United States; in 2008-2010, cases = were=20 reported in every state except Alaska and Hawaii, and Puerto Rico.

Rabid bats may be clumsy, disoriented, and unable to fly, which makes = it more=20 likely they will come into contact with humans. One should avoid = handling them=20 or having them in one's living space, as with any wild animal. If a bat = is found=20 in living quarters near a child, mentally handicapped person, = intoxicated=20 person, sleeping person, or pet, living in an area where rabies is known = to=20 occur, the person or pet should receive immediate medical attention for = rabies.=20 Bats have very small teeth and can bite a sleeping person without being = felt.=20 There is evidence that it is possible for the bat rabies virus to infect = victims=20 purely through airborne transmission, without direct physical contact of = the=20 victim with the bat itself.[83][84]

If a bat is found in a house and the possibility of exposure cannot = be ruled=20 out, the bat should be sequestered and an animal control officer called=20 immediately, so the bat can be analysed. This also applies if the bat is = found=20 dead. If it is certain nobody has been exposed to the bat, it should be = removed=20 from the house. The best way to do this with live bats is to close all = the doors=20 and windows to the room except one that opens to the outside. The bat = should=20 soon leave.

Due to the risk of rabies and to health problems related to their = faecal=20 droppings, bats should be excluded from inhabited parts of houses. The = Centers=20 for Disease Control and Prevention (CDC) provide fullly detailed = information=20 on all aspects of bat management, including how to capture a bat, what = to do in=20 case of exposure, and how to bat-proof a house humanely.[85]=20 In certain countries, such as the United=20 Kingdom, it is illegal to handle bats without a license.

Where rabies is not endemic, as throughout most of Western Europe, = small=20 bats can be considered harmless. Larger bats can give a nasty bite.

[edit]=20 Mythology

In European cultures, bats have long been associated with witchcraft, = black=20 magic and darkness. The witches incorporate bat in their brew in = Shakespeare's=20 Macbeth.[<= /SPAN>86]=20 Because bats are mammals, yet can fly, this gives them status as liminal beings = in many=20 cultural traditions.[citation=20 needed]

The bat is sacred in Tonga=20 and is often considered the physical manifestation of a separable soul.[87]=20 Bats are closely associated with vampires,=20 who are said to be able to shapeshift=20 into bats, fog, or wolves. Bats are also = symbols of ghosts, = death, and disease. Among some Native=20 Americans, such as the Creek,=20 Cherokee=20 and Apache,=20 the bat is a trickster = spirit.

Chinese lore=20 claims the bat is a symbol of longevity and happiness, and is similarly = lucky in=20 Poland and=20 geographical Macedonia=20 and among the Kwakiutl and Arabs.

Pre-Columbian=20 cultures associated animals with gods, and often displayed them in art. = The Moche = people=20 depicted bats in their ceramics.[88]

In Western=20 culture, the bat is often a symbol of the night and its foreboding = nature.=20 The bat is a primary animal associated with fictional characters of the = night,=20 both villains, such as Dracula, and heroes, such as Batman. The association = of the=20 fear of the night with the animal was treated as a literary challenge by = Kenneth=20 Oppel, who created a best-selling series of novels, beginning with = Silverwing,=20 which feature bats as the central heroic figures much as = anthropomorphized rabbits were the=20 central figures to the classic novel=20 Watership = Down.

An old=20 wives' tale has it that bats will entangle themselves in people's = hair. One=20 likely source of this belief is that insect-eating bats seeking prey may = dive=20 erratically toward people, who attract mosquitoes=20 and gnats, leading=20 the squeamish to believe the bats are trying to get in their hair.

[edit]=20 Mesoamerica

In Mesoamerican mythology during the = Classic-Contemporary period, bats symbolized the land of the dead, which = was=20 considered to be the underworld.[89]=20 They also symbolized destruction and decay. Bats may have symbolized in = this way=20 because they fly only at night and dwell in caves during the daytime and = are=20 associated with human skulls and bones by classic Maya = ceramists.=20 Central Mexicans sometimes depicted bats having snouts that looked like=20 "sacrificial knives and carrying human head" in the Postclassic era.[90]=20 Bat images were engraved onto funerary urns, and were emphasized with = large=20 claws and round ears by Zapotecs. = They were commonly associated with death.[91]=20 The depiction of bats on funeral urns and goods took on some the = characteristics=20 of the jaguar,=20 which was, and still is, another entity of the night and the underworld. = There=20 have also been instances where bats are portrayed next to other animals=20 portrayed negatively in Mesoamerica, including scorpions=20 and other nocturnal animals such as owls.

A gigantic, life-size, ceramic bat-man has been discovered and dug up = from=20 the Templo Mayor.[citation=20 needed] The Templo Mayor is located in the center = of the Mexica = capital of=20 Tenochtitlan. = Known as a=20 god of death, this statue has the clawed feet and hands of a bat, but = the body=20 of a man. The statue's human-like eyes bulged out from the bat-like = head, making=20 the Zapotec=20 images very realistic and living. In the 1930s, the Kaqchikel = Maya were=20 said to have proclaimed the bat was the Devil=E2=80=99s provider. = Kaqchikel would leave=20 the Devil=E2=80=99s underworld home and collect blood from the animals = to be used for=20 scrumptious meals to feed the Devil. =E2=80=9CIn the myths, the beast of = prey and the=20 animal that is preyed upon play two significant roles. They represent = two=20 aspects of life=E2=80=94the aggressive, killing, conquering, creating = aspect of life,=20 and the one that is the matter or, you might say, the subject = matter=E2=80=9D.[92]=20 In the Devil=E2=80=99s underworld, dead sinners would work off their = sins to get to=20 heaven, indicating the bat, too, was a sinner and worked under the = authority of=20 the Devil.[93]

[edit]=20 Oaxaca

Oaxacans=20 believe the jealousy of the bat in wanting birds' feathers that gently = fit their=20 bodies led him to become nocturnal. The bat feeling isolated and = undesirable=20 spoke to God after that he complained he was extremely cold. God, fair = and just,=20 turned to birds in the animal kingdom and asked if they would show = compassion=20 and donate a feather to the bat so the feathers would keep him warm. The = birds=20 all agreed, and began to pluck one feather from their bodies to give to = the bat.=20 With all of the feathers, the bat became more magnificent-looking than = all=20 birds, even able to spread color to the night sky. During daylight, the = bat=20 created rainbows that reflected vibrant colors from the sun. The bat = soon became=20 overly arrogant and conceited, due having this new and improved look. = The birds=20 grew tired of the bat=E2=80=99s self conceitedness and glorification, = and so decided to=20 fly up to heaven and speak to God to do something. The birds informed = God of the=20 bat's behaviour, God was surprised and so decided to take a look = himself. When=20 on Earth, God called on the bat to show him what he was doing. The bat = began to=20 fly across the light blue sky, where one by one each feather began to = fall out,=20 uncovering the bat=E2=80=99s natural, ugly-looking body. The bat became = ashamed and=20 distressed of his appearance after all feathers came off, missing the = beautiful,=20 plentiful feathers that he had, so he decided to hide in caves during = the day.=20 He would only come out during the night, searching high and low for the = feathers=20 to avoid embarrassment that he will not be seen during his search.[93]

[edit]=20 East Nigeria

According to a particular East Nigerian tale, the bat developed its = nocturnal=20 habits after causing the death of his partner the bush-rat. The bat and = the=20 bush-rat would share activities such as rummaging through the grass and = trees,=20 hunting, talking and bonding during the day. When at night, the bat and = the=20 bush-rat would alternate in cooking duties, cooking what was caught, and = eat=20 together. It appeared to be a dedicated partnership, but the bat hated = the=20 bush-rat immensely. The bush-rat always found the bat=E2=80=99s soup=20 more appetising, so when eating dinner one night, asked the bat why the = soup=20 tasted better than his own and also asked how it was made. The bat = agreed to=20 show him how to make it the next day, but instead was forming a = malicious=20 plan.

Next day as bat prepared his soup, the bush-rat came, greeting him = and asking=20 if he could be shown what was agreed yesterday. Earlier, the bat has = found a pot=20 looking exactly like the one he used usually, but it held warm water and = so=20 decided to use this instead. The bat explained to the bush-rat that to = make his=20 soup, he had to boil himself prior to serving the soup, where sweetness = and=20 flavor of the soup came from the flesh.=20 The bat jumped in the pot seemingly excited, with the bush-rat = mesmerised. After=20 a few minutes the bat climbed out and while the bush-rat was distracted, = switched pots. The bat then served his soup out of the soup pot, both = tasted it.=20 Overanxious and eager, the bush-rat jumped into the pot of warm water. = He stayed=20 much longer in the pot, dying in the process.

When the bush-rat=E2=80=99s wife returned that night = to find=20 her husband dead, she wept and ran to the chief=20 of the land's house, telling him about what happened and what she was = sure what=20 the bat had done. In hearing this, the chief became angry, ordering for = the=20 immediate arrest of the bat. It just so happened that the bat was flying = over=20 the house and overheard what was just said. He quickly went into hiding = high up=20 in a tree. When the chief=E2=80=99s men went looking for the bat, he = could not be found.=20 The search to arrest the bat carried on over several days, but he still = could=20 not be found. The bat needed to eat, so flew out of hiding every night = to hunt = for food to = escape of=20 being arrested. This, according to Eastern Nigeria mythology, is why = bats only=20 fly at night.[94]

[edit]=20 Heraldry

Main article: Bat = (heraldry)
Burgee of the Royal Valencia Yacht = Club

The bat is sometimes used as a heraldic=20 symbol. The coats of arms of certain cities in eastern Spain, such as Valencia, Palma de = Mallorca and=20 Fraga, have the=20 bat over the shield. Formerly, the Barcelona=20 city coat of arms also had a bat crowning it, but the bat has been = removed in=20 the present-day versions.

The heraldic use of the bat in Valencia, Catalonia=20 and the Balearic=20 Islands has its origins in a winged dragon (vibra=20 or vibria), which featured in King=20 James I of Aragon's helmet or ci= mera=20 reial. This is the most widely accepted theory, although there = is also a=20 legend that says that due to the intervention of a bat, King James was = able to=20 win a crucial battle against the Saracens=20 that allowed him to win Valencia for his kingdom.

The use of the bat as a heraldic symbol is prevalent in the = territories of=20 the former Crown=20 of Aragon and it is little used elsewhere. However, it can be found = in a few=20 places, as in the coats of arms of the city of Albacete,=20 in Spain, as well as the town of Montchauvet=20 (Yvelines), in France.

Certain Spanish football=20 clubs including Valencia=20 CF and Levante UD use bats = in their=20 badges.

The Burgee=20 of the Royal Valencia Yacht Club (Reial Club N=C3=A0utic de = Val=C3=A8ncia) displays a bat=20 on a golden field in its center.

[edit]=20 State = symbols

Three U.S. states have an official state=20 bat. Texas and Oklahoma are represented by the Mexican = free-tailed=20 bat; Virginia is represented by the Virginia = big-eared=20 bat.[95]

[edit]=20 See also

[edit]=20 References

Notes
  1. ^ a=20 b=20 Eick = et al.;=20 Jacobs, DS; Matthee, CA (2005). "A Nuclear DNA Phylogenetic = Perspective on the=20 Evolution of Echolocation and Historical Biogeography of Extant Bats=20 (Chiroptera)". Molecular Biology and Evolution 22 (9): = 1869. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1093%2Fmolbev%2Fmsi180" = rel=3D"nofollow">10.1093/molbev/msi180.=20 PMID 15930153.=20 "Several molecular studies have shown that Chiroptera belong to the=20 Laurasiatheria (represented by carnivores, pangolins, cetartiodactyls, = eulipotyphlans, and perissodactyls) and are only distantly related to=20 dermopterans, scandentians, and primates (Nikaido et al. 2000; Lin and = Penny=20 2001; Madsen et al. 2001; Murphy et al. 2001a, 2001b; = Van Den=20 Bussche and Hoofer 2004)." 
  2. ^=20 =CF=87=CE=B5=CE=AF=CF=81, Henry George Liddell, = Robert Scott, A=20 Greek-English Lexicon, on Perseus
  3. ^=20 =CF=80=CF=84=CE=B5=CF=81=CF=8C=CE=BD, Henry = George Liddell, Robert Scott, A=20 Greek-English Lexicon, on Perseus
  4. ^ <= B>a=20 <= B>b=20 Hunter, P.=20 (September 2007). "The nature of flight. The molecules and mechanics of = flight in=20 animals". Science and Society 8 (9): = 811=E2=80=93813. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1038%2Fsj.embor.7401050" = rel=3D"nofollow">10.1038/sj.embor.7401050.=20 PMC 1973956. PMID 17767190. http://ukpmc.ac.uk/articlerender.cgi?artid=3D1635883= . Retrieved = 2009-07-17. 
  5. ^=20 Tudge, = Colin (2000).=20 The Variety of Life. Oxford University Press. = ISBN 0= -19-860426-2. 
  6. ^=20 "Bumblebee=20 bat (Craseonycteris thonglongyai)". EDGE=20 Species. http://www.edgeofexistence.org/mammals/species_info.php?= id=3D49. Retrieved = 2008-04-10. 
  7. ^=20 Kitti's Hog-Nosed Bat: Craseonycteridae - Physical=20 Characteristics - Bats, Bumblebee, Species, Inches, Brown, and=20 Tips
  8. ^=20 Wood, = Gerald (1983).=20 The Guinness Book of Animal Facts and Feats. = ISBN 978-0-85112-235-9. 
  9. ^=20 ARKive - Golden-capped fruit bat videos, photos and = facts -=20 Acerodon jubatus
  10. ^=20 Online Etymology Dictionary: flitter = (verb)
  11. ^ = = a=20 = b=20 Nancy = B. Simmons1,=20 Kevin L. Seymour2, Jo=C2=A8rg Habersetzer3 & Gregg F. Gunnell4 = (2008).=20 "Primitive Early Eocene bat from Wyoming and the evolution of flight = and=20 echolocation". Nature 451 (7180): 818. Bibcode=20 2008Natur.451..818S. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1038%2Fnature06549" = rel=3D"nofollow">10.1038/nature06549.=20 PMID 18270539. 
  12. ^=20 Teeling et=20 al. (2005). "A Molecular Phylogeny for Bats Illuminates = Biogeography and=20 the Fossil Record". Science 307: 580=E2=80=93584. Bibcode 2005Sci...307..580T. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1126%2Fscience.1105113" = rel=3D"nofollow">10.1126/science.1105113.=20 PMID 15681385. 
  13. ^=20 Teeling et=20 al. (2000). "Molecular evidence regarding the origin of = echolocation and=20 flight in bats". Nature 403 (6766): 188=E2=80=93192. Bibcode 2000Natur.403..188T. doi:<= A=20 class=3D"external text" href=3D"http://dx.doi.org/10.1038%2F35003188" = rel=3D"nofollow">10.1038/35003188. 
  14. ^=20 "Order Chiroptera (Bats)". Animal Diversity = Web.=20 http://animaldiversity.ummz.umich.edu/site/accounts/info= rmation/Chiroptera.html. Retrieved = 2007-12-30. 
  15. ^=20 Springer et=20 al. (2001). "Integrated fossil and molecular data reconstruct bat = echolocation". Proceedings of the National Academy of = Sciences=20 98: 6241=E2=80=936246. Bibcode=20 2001PNAS...98.6241S. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1073%2Fpnas.111551998" = rel=3D"nofollow">10.1073/pnas.111551998.=20 PMC 33452.=20 PMID 11353869.=20 = //www.pubmedcentral.nih.gov/articlerender.fcgi?tool=3Dpmcentrez&artid= =3D33452. 
  16. ^=20 Li et al. 2007. "Accelerated FoxP2 = Evolution in=20 Echolocating Bats". PLOS ONE.
  17. ^=20 Li et al. 2008. "The hearing gene = Prestin=20 reunites the echolocating bats". Proc. Natl. Academy. Sci. = U.S.A.
  18. ^= =20 James M. = Hutcheon and=20 John A.W. Kirsch. "A moveable face: deconstructing the Microchiroptera = and a new=20 classification of extant bats" (PDF). doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.3161%2F1733-5329%282006%298%255B1%3AAMFDTM%2= 55D2.0.CO%3B2"=20 = rel=3D"nofollow">10.3161/1733-5329(2006)8%5B1:AMFDTM%5D2.0.CO;2.=20 http://www.bio.georgiasouthern.edu/bio-home/hutcheon/hut= chkirsch2006.pdf. [dead=20 link]
  19. ^=20 Eick = et al.;=20 Jacobs, DS; Matthee, CA (2005). "A Nuclear DNA Phylogenetic = Perspective on the=20 Evolution of Echolocation and Historical Biogeography of Extant Bats=20 (Chiroptera)". Molecular Biology and Evolution 22 (9): = 1869. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1093%2Fmolbev%2Fmsi180" = rel=3D"nofollow">10.1093/molbev/msi180.=20 PMID 15930153.=20 "Following the recommendations of Hutcheon and Kirsch (2004), we refer = to the=20 two suborders of chiropterans as =E2=80=98Pteropodiformes=E2=80=99 = (comprising the=20 Pteropodidae, Rhinolophidae, Hipposideridae, Megadermatidae, and=20 Rhinopomatidae) and =E2=80=98Vespertilioniformes=E2=80=99 (remaining = microbat=20 families)." 
  20. ^=20 "Primitive Early Eocene bat from Wyoming and the = evolution of=20 flight and echolocation". Nature. 2008-02-14. http://www.nature.com/nature/journal/v451/n7180/abs/natu= re06549.html. Retrieved = 2008-07-03. 
  21. ^=20 Pettigrew JD,=20 Maseko BC, Manger PR (April 2008). "Primate-like retinotectal = decussation in=20 an echolocating megabat, Rousettus aegyptiacus". Neuroscience=20 153 (1): 226=E2=80=9331. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1016%2Fj.neuroscience.2008.02.019"=20 rel=3D"nofollow">10.1016/j.neuroscience.2008.02.019. PMID 18367343. 
  22. ^=20 Eiting, T.P. and G.F. Gunnell. 2009. = Global=20 completeness of the bat fossil record. Journal of Mammalian Evolution=20 16:151-173.
  23. ^=20 Simmons, N.B., K.L. Seymour, J. = Habersetzer, and=20 G.F. Gunnell. 2008. Primitive early Eocene bat from Wyoming and the = evolution=20 of flight and echolocation. Nature 451:818-U816.
  24. ^=20 (BBC News) "Bat fossil solves evolution = poser" 13=20 February 2008.
  25. ^=20 (Discovery Channel article) "Prehistoric bats = learned to=20 fly before they could see" February 13, 2008.[dead=20 link]
  26. ^=20 Thomas, S.P., and R.A. Suthers. 1972. = Physiology=20 and energetics of bat flight. Journal of Experimental Biology=20 57:317-&.
  27. ^=20 "Bats of the World". Bat Conservation Trust.=20 http://www.bats.org.uk/pages/bats_of_the_world.html<= /SPAN>. Retrieved January = 2011. 
  28. ^=20 Grzimek's Animal=20 Life Encyclopedia: Vol 13 Mammals II (2nd ed.). 2003. p. 311. = = ISBN 0= -7876-5362-4. 
  29. ^=20 "Bat Conservation International: Species = Profiles". http://www.batcon.org/index.php/all-about-bats/species-p= rofiles.html. 
  30. ^=20 "The Mysterious White Nose Syndrome and the 10 Bat = Species Most=20 at Risk". http://www.thedailygreen.com/environmental-news/latest/w= hite-nose-syndrome-in-bats-0930. 
  31. ^=20 "California=20 Bats". http://www.batcrew.com/cabats.html. 
  32. ^=20 http://www.avonbatgroup.org.uk/resources/Wavesurfer+inst= ructions-draft.pdf=20 page 10
  33. ^=20 Jones, = K. E., O.=20 R. P. Bininda-Emonds, and J. L. Gittleman (2005). "Bats, clocks, and = rocks:=20 diversification patterns in chiroptera". Evolution 59 = (10):=20 2243=E2=80=932255. PMID 16405167. 
  34. ^=20 Teeling, E. C.=20 (2009). "Hear, hear: the convergent evolution of echolocation in = bats?".=20 Trends in Ecology & Evolution 24 (7): = 351=E2=80=93354. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1016%2FJ.Tree.2009.02.012"=20 rel=3D"nofollow">10.1016/J.Tree.2009.02.012. PMID 19482373. 
  35. ^=20 Jones, = G., and M.=20 W. Holderied (2007). "Bat echolocation calls: adaptation and convergent=20 evolution". Proceedings of the Royal Society B-Biological = Sciences=20 274 (1612): 905=E2=80=93912. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1098%2FRspb.2006.0200" = rel=3D"nofollow">10.1098/Rspb.2006.0200.=20 PMC 1919403. PMID 17251105.=20 = //www.pubmedcentral.nih.gov/articlerender.fcgi?tool=3Dpmcentrez&artid= =3D1919403. 
  36. ^=20 DesRoche, K., M.=20 B. Fenton, and W. C. Lancaster (2007). "Echolocation and the thoracic=20 skeletons of bats: a comparative morphological study". Acta=20 Chiropterologica 9 (2): 483=E2=80=93494. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.3161%2F1733-5329%282007%299%5B483%3AEATTSO%5= D2.0.CO%3B2"=20 rel=3D"nofollow">10.3161/1733-5329(2007)9[483:EATTSO]2.0.CO;2. ISSN 1733-5329. 
  37. ^=20 http://www.jstor.org/pss/3546476
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  41. ^ = a=20 = b=20 Roberts, W.C.=20 (October 2006). "Facts and ideas from anywhere". Proceedings = (Baylor=20 University. Medical Center) 19 (4): 425=E2=80=93434. PMC 1618737. PMID 17106509. http://ukpmc.ac.uk/articlerender.cgi?artid=3D804041<= /SPAN>. Retrieved = 2009-07-17. 
  42. ^=20 Irwin, = N. (March=20 1997). "Wanted DNA samples from Nyctimene or = Paranyctimene=20 Bats" (PDF). The New Guinea Tropical Ecology and = Biodiversity=20 Digest 3: 10. http://papuaweb.anu.edu.au/dlib/jr/ngtebd/03.pdf. Retrieved = 2009-07-17. 
  43. ^ = = a=20 = b=20 = c=20 Melissa = Calhoun (15=20 December 2005). "Bats Use Touch Receptors on Wings to Fly, Catch = Prey, Study=20 Finds". http://news.research.ohiou.edu/news/index.php?item=3D257= . Retrieved = 2006-10-18. 
  44. ^=20 Chamberlain, Ted=20 (2006-12-06). "Photo in the News: Bat Has Longest Tongue of Any = Mammal".=20 National Geographic News. National Geographic Society.=20 http://news.nationalgeographic.com/news/2006/12/061206-t= ongue-photo.html. Retrieved 2007-06-18. "A.=20 fistulata (shown lapping sugar water from a tube) has the longest = tongue,=20 relative to body length, of any mammal=E2=80=94and now scientists = think they know=20 why." 
  45. ^=20 Smithsonian Institute
  46. ^=20 Fenton, = M. Brock=20 (2001). Bats. New York: Checkmark Books. = pp. 60=E2=80=9362. = ISBN 0= -8160-4358-2. 
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  48. ^ a= =20 b= =20 Fenton, = M. Brock=20 (2001). Bats. New York: Checkmark Books. = pp. 95=E2=80=93107. = ISBN 0= -8160-4358-2. 
  49. ^ a= =20 b= =20 Fenton, = M. Brock=20 (2001). Bats. New York: Checkmark Books. pp. 4=E2=80=935. = = ISBN 0= -8160-4358-2. 
  50. ^=20 "Biology of Bats". http://books.google.co.nz/books?id=3DGtp4yWnPD9YC&pr= intsec=3Dfrontcover&dq=3DBiology+Bats&hl=3Den&sa=3DX&ei=3D= -erJT5mvLO2UiAeD6IngBg&ved=3D0CDUQ6AEwAA#v=3Donepage&q=3DBiology%= 20Bats&f=3Dfalse. 
  51. ^=20 Retrieved 22 October 2006.[dead=20 link]
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  56. ^=20 A trace of GLO was detected in only one = of 34 bat=20 species tested, across the range of six families of bats tested: See = Jenness,=20 R., E. Birney, and K. Ayaz. 1980. Variation of L-gulonolactone oxidase = activity in placental mammals. Comparative Biochemistry and Physiology = 67B:195-204. Earlier reports of only fruit bats being deficient were = based on=20 smaller samples.
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  58. ^=20 Arita, Hector T., and Karina = Santos-del-Prado.=20 "Conservation Biology of Nectar-Feeding Bats in Mexico." Journal of = Mammalogy=20 80.1 (1999): 31=E2=80=9341
  59. ^=20 M, L. Gerardo H., Keith A. Hobson, = Adriana M. A,=20 Daniel E. B, Victor Sanchez-Corero, and German M. C. "The Role of = Fruits and=20 Insects in the Nutrition of Frugivorous Bats: Evaluating the Use of = Stable=20 Isotope Models." Biotropica 33.3 (2001): 520=E2=80=9328
  60. ^=20 Greenhall, Arthur M. 1961. Bats in = Agriculture. A=20 Ministry of Agriculture Publication. Trinidad and Tobago
  61. ^=20 Jones, V. = (2000). "Drinking=20 in the river". Vivian Jones. http://www.bellingen.com/flyingfoxes.=20 Retrieved 17 July 2009. 
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  64. ^=20 "Protecting and managing underground sites for bats = (pdf), see=20 section 6.4." (PDF). http://www.eurobats.org/documents/pdf/AC9/Doc_AC9_15_Pro= tecting_underground_sites.pdf. Retrieved = 2006-05-18. 
  65. ^=20 "Pillbox=20 converted to bat retreat, BBC website". BBC News. = 2006-04-06. http://news.bbc.co.uk/1/hi/england/4885642.stm. Retrieved = 2006-05-18. 
  66. ^= =20 "Fish and Wildlife Service Awards $800,000 in Grants = to Explore=20 Cause, Control of White-Nose Syndrome in Bats". United=20 States Fish and Wildlife Service. 2009-10-26. http://www.fws.gov/news/NewsReleases/showNews.cfm?newsId= =3D9191BAAD-F8E8-0097-B3670BDF3849EBF2. Retrieved = 2009-10-30. 
  67. ^ a= =20 b= =20 c= =20 "Culprit=20 Identified: Fungus Causes Deadly Bat Disease". Uni= ted=20 States Geological Survey. 2011-10-26. http://www.usgs.gov/newsroom/article.asp?ID=3D3015#.T86q= bVJ42uK. Retrieved = 2012-5-5. 
  68. ^=20 "White-nose syndrome in bats =E2=80=93 Frequently = Asked Questions"=20 (PDF). United=20 States Fish and Wildlife Service. 2009-04. http://www.fws.gov/northeast/pdf/white-nosefaqs.pdf<= /SPAN>. Retrieved = 2009-10-27. 
  69. ^=20 p63 Safe= ty and=20 Health Practitioner February 2010
  70. ^=20 Erin F. Baerwald et al, Barotrauma is a = significant cause of bat fatalities at wind turbines Current = Biology,=20 2008
  71. ^ a<= /B>=20 b<= /B>=20 "B.C. study to help bats survive wind farms[dead=20 link]", National=20 Wind Watch, September 23, 2008
  72. ^=20 "Bats take a battering at wind farms", New = Scientist, May=20 12, 2007
  73. ^=20 "Caution Regarding Placement of Wind Turbines on = Wooded Ridge=20 Tops" (PDF). Bat Conservation International. 4 January 2005.=20 http://vawind.org/Assets/Docs/BCI_ridgetop_advisory.pdf<= /A>. Retrieved = 2006-04-21. 
  74. ^=20 Arnett, = Edward B.;=20 Wallace P. Erickson, Jessica Kerns, Jason Horn (June 2005). "Relationships between Bats and Wind Turbines in = Pennsylvania=20 and West Virginia: An Assessment of Fatality Search Protocols, = Patterns of=20 Fatality, and Behavioral Interactions with Wind Turbines" (PDF). = Bat=20 Conservation International. Archived from the original on 2006-02-10. http://web.archive.org/web/20060210183113/http://batcon.= org/wind/BWEC2004finalreport.pdf. Retrieved = 2006-04-21. 
  75. ^=20 Baerwald, Erin F;=20 D'Amours, Genevieve H; Klug, Brandon J; Barclay, Robert MR = (2008-08-26).=20 "Barotrauma is a significant cause of bat fatalities at wind = turbines". Current = Biology=20 18 (16): R695=E2=80=93R696. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1016%2Fj.cub.2008.06.029"=20 rel=3D"nofollow">10.1016/j.cub.2008.06.029. OCLC=  252616082.=20 PMID 18727900.=20 Lay summary =E2=80=93 CBC=20 Radio =E2=80=93 Quirks=20 & Quarks (2008-09-20). [dead=20 link] Laysource includes audio podcast of = interview with=20 author.
  76. ^=20 Wong, = Samson;=20 Susanna Lau, Patrick Woo, Kwok-Yung Yuen (2006-10-16). "Bats as a continuing source of emerging infections = in=20 humans" (Review). Reviews=20 in Medical Virology (John=20 Wiley & Sons) 17 (2): 67=E2=80=9391. doi:<= A=20 class=3D"external text" href=3D"http://dx.doi.org/10.1002%2Frmv.520" = rel=3D"nofollow">10.1002/rmv.520.=20 PMID 17042030. http://www3.interscience.wiley.com/cgi-bin/abstract/1133= 98566/ABSTRACT?CRETRY=3D1&SRETRY=3D0. Retrieved 2007-12-29. "The = currently=20 known viruses that have been found in bats are reviewed and the risks = of=20 transmission to humans are highlighted. (from abstract)" 
  77. ^<= /B>=20 McColl, KA; N=20 Tordo, AA Aquilar Setien (April 2000). "Bat lyssavirus infections". = Revue=20 scientifique et technique 19 (1): 177=E2=80=93196. PMID 11189715.=20 "Bats, which represent approximately 24% of all known mammalian = species,=20 frequently act as vectors of lyssaviruses. (from = abstract)" 
  78. ^<= /SPAN>=20 Li, = Wendong; Z.=20 Shi, M. Yu, W. Ren and 13 additional coauthors (2005-10-28). "Bats=20 are natural reservoirs of SARS-like coronaviruses". Science=20 310 (5748): 676=E2=80=93679. Bibcode=20 2005Sci...310..676L. doi:<= A=20 class=3D"external text" = href=3D"http://dx.doi.org/10.1126%2Fscience.1118391" = rel=3D"nofollow">10.1126/science.1118391.=20 PMID 16195424. http://www.sciencemag.org/cgi/content/abstract/310/5748/= 676. Retrieved 2007-12-29. Lay=20 summary =E2=80=93 Science (2005-10-28). "The = genetic diversity of=20 bat-derived sequences supports the notion that bats are a natural = reservoir=20 host of the SARS cluster of coronaviruses." 
  79. ^<= /B>=20 Halpin, K.; P. L.=20 Young, H. E. Field and J. S. Mackenzie (August 1, 2000). "Isolation=20 of Hendra virus from pteropid bats: a natural reservoir of Hendra = virus".=20 Journal= =20 of General Virology 81 (8): 1927=E2=80=931932. PMID 10900029. http://vir.sgmjournals.org/cgi/content/abstract/81/8/192= 7. Retrieved 2007-12-29. "In this = paper we=20 describe the isolation of HeV from pteropid bats, corroborating our=20 serological and epidemiological evidence that these animals are a = natural=20 reservoir host of this virus." 
  80. ^=20 Leroy, = Eric M.;=20 Brice Kumulungui, Xavier Pourrut, Pierre Rouquet and 6 additional = coauthors=20 (2005-12-01). "Fruit bats as reservoirs of Ebola virus" (Brief=20 Communication). Nature=20 438 (7068): 575=E2=80=93576. Bibcode=20 2005Natur.438..575L. doi:<= A=20 class=3D"external text" href=3D"http://dx.doi.org/10.1038%2F438575a" = rel=3D"nofollow">10.1038/438575a.=20 PMID 16319873. http://www.nature.com/nature/journal/v438/n7068/abs/4385= 75a.html. Retrieved 2007-12-29. "We find = evidence=20 of asymptomatic infection by Ebola virus in three species of fruit = bat,=20 indicating that these animals may be acting as a reservoir for this = deadly=20 virus. (from abstract)" 
  81. ^=20 Charles = Q. Choi=20 (March 2006). "Going to Bat". Scientific=20 American: pp. 24, 26. http://www.sciam.com/article.cfm?id=3Dgoing-to-bat. Retrieved 2007-12-29. "Long = known as=20 vectors for rabies, bats may be the origin of some of the most deadly = emerging=20 viruses, including SARS, Ebola, Nipah, Hendra and = Marburg."  Note: This could be = considered a=20 lay summary of the various scientific publications cited in the = preceding=20 sentence.
  82. ^=20 "Center for Disease Control and Prevention: Human=20 Rabies". http://www.cdc.gov/rabies/location/usa/surveillance/huma= n_rabies.html. 
  83. ^=20 Constantine, Denny=20 G. (April 1962). "Rabies transmission by nonbite route". Public=20 Health Reports (Public=20 Health Service) 77 (4): 287=E2=80=93289. PMC 1914752. PMID 13880956.=20 = //www.pubmedcentral.nih.gov/articlerender.fcgi?tool=3Dpmcentrez&artid= =3D1914752. Retrieved 2007-12-29. "These = findings=20 support consideration of an airborne medium, such as an aerosol, as = the=20 mechanism of rabies transmission in this instance." 
  84. ^=20 Messenger, Sharon=20 L.; Jean S. Smith and Charles E. Rupprecht (2002-09-15). "Emerging Epidemiology of Bat-Associated Cryptic = Cases of=20 Rabies in Humans in the United States". Clinic= al=20 Infectious Diseases 35 (6): 738=E2=80=93747. doi:<= A=20 class=3D"external text" href=3D"http://dx.doi.org/10.1086%2F342387"=20 rel=3D"nofollow">10.1086/342387. PMID 12203172. http://www.journals.uchicago.edu/doi/abs/10.1086/342387<= /A>. Retrieved 2007-12-29. "Cryptic = rabies=20 cases are those in which a clear history of exposure to rabies virus = cannot be=20 documented, despite extensive case=E2=80=90history investigation. = Absence of a=20 documented bite history reflects inherent difficulties in obtaining = accurate=20 animal=E2=80=90contact information.... <gap> Thus, absence of = bite=E2=80=90history data=20 does not mean that a bite did not occur." 
  85. ^=20 CDC's website on bats and rabies
  86. ^=20 de Vries, = Ad (1976).=20 Dictionary of Symbols and Imagery. Amsterdam: North-Holland = Publishing=20 Company. p. 36. = ISBN 0= -7204-8021-3. 
  87. ^=20 Kingdom of Tonga: Safe Haven for Flying = Foxes
  88. ^=20 Berrin, Katherine & Larco Museum. = The=20 Spirit of Ancient Peru:Treasures from the Museo=20 Arqueol=C3=B3gico Rafael Larco Herrera. New York: Thames=20 and Hudson, 1997.
  89. ^=20 Aztec Symbols
  90. ^=20 Kay Almere Read and Jason J. Gonzalez. = 2000.=20 Mesoamerican Mythology. Oxford University Press. pp. 132
  91. ^=20 Artists Inspired by Oaxaca Folklore Myths and=20 Legends
  92. ^=20 Joseph Cambell and Bill Moyers. 1988. = The Power=20 of Myth. Doubleday. pp. 91
  93. ^=20 a=20 b=20 Kay Almere Read and Jason J. Gonzalez. = 2000.=20 Mesoamerican Mythology. Oxford University Press. pp. = 132=E2=80=93134
  94. ^=20 Arnott, Kathleen. 1962. African Myths = and=20 Legends. Oxford University Press. Pp. 150=E2=80=93152
  95. ^=20 "Official state=20 bats". netstate.com. NSTATE, LLC.. Archived from the=20 original on March 8, 2008. http://www.webcitation.org/5x2xEXoK2.=20 Retrieved February 13, 2011. 
Bibliography
  • Greenhall, Arthur H. 1961. Bats in Agriculture. A Ministry = of=20 Agriculture Publication. Trinidad and Tobago.
  • Nowak, Ronald M. 1994. " Walker's BATS of the World". The Johns = Hopikins=20 University Press, Baltimore and London.
  • John D. Pettigrew's summary on Flying Primate=20 Hypothesis
  • Altringham, J.D. 1998. Bats: Biology and Behaviour. Oxford: Oxford = University Press.
  • Dobat, K.; Holle, T.P. 1985. Bl=C3=BCten und Flederm=C3=A4use: = Best=C3=A4ubung durch=20 Flederm=C3=A4use und Flughunde (Chiropterophilie). Frankfurt am Main: = W. Kramer=20 & Co. Druckerei.
  • Fenton, M.B. 1985. Communication in the Chiroptera. Bloomington: = Indiana=20 University Press.
  • Findley, J.S. 1995. Bats: a Community Perspective. Cambridge: = Press=20 Syndicate of the University of Cambridge.
  • Fleming, T.H. 1988. The Short-Tailed Fruit Bat: a Study in = Plant-Animal=20 Interactions. Chicago: The University of Chicago Press.
  • Kunz, T.H. 1982. Ecology of Bats. New York: Plenum Press.
  • Kunz, T.H.; Racey, P.A. 1999. Bat Biology and Conservation. = Washington:=20 Smithsonian Institution Press.
  • Kunz, T.H.; Fenton, M.B. 2003. Bat Ecology. Chicago: The = University of=20 Chicago Press.
  • Neuweiler, G. 1993. Biologie der Flederm=C3=A4use. Stuttgart: = Georg Thieme=20 Verlag.
  • Nowak, R.M. 1994. Walker's Bats of the World. Baltimore: The Johns = Hopkins=20 University Press.
  • Richarz, K. & Limbruner, A. 1993. The World of Bats. Neptune = City: TFH=20 Publications.
  • Teeling, E.C. 2009. Chiroptera. Oxford University Press.
  • Twilton, B. 1999. My Life as The Bat. Liverpool Hope University = press
Further reading

[edit]=20 External = links

Find = more about=20 bat on Wikipedia's sister=20 projects:
Definitions=20 and translations from Wiktionary
Images=20 and media from Commons
Learning=20 resources from Wikiversity
News=20 stories from Wikinews
Quotations=20 from Wikiquote
Source=20 texts from Wikisource
Textbooks=20 from Wikibooks
3D"" Wikispecies=20 has information related to: Chiroptera<= /I>
3D"" The Wikibook Dichotomous=20 Key has a page on the topic of
=
Extant mammal=20 orders by infraclass
Australosphenida
Metatheria
(Marsupial=20 inclusive)
=
Eutheria
(Placental=20 inclusive)
Chiroptera = families
Yinpterochiropte= ra
(Pteropodiformes)
Yangochiroptera
= (Vespertilioniformes)

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.toctitle { text-align: center; } #toc ul { padding: 0px; text-align: left; margin-left: 0px; list-style-type: = none; list-style-image: none; } .toc ul { padding: 0px; text-align: left; margin-left: 0px; list-style-type: = none; list-style-image: none; } #toc ul ul { margin: 0px 0px 0px 2em; } .toc ul ul { margin: 0px 0px 0px 2em; } #toc .toctoggle { font-size: 94%; } .toc .toctoggle { font-size: 94%; } .toccolours { padding: 5px; border: 1px solid rgb(170, 170, 170); font-size: 95%; = background-color: rgb(249, 249, 249); } .mw-warning { text-align: center; margin-right: 50px; margin-left: 50px; } div.floatright { margin: 0px 0px 0.5em 0.5em; border: 0px currentColor; } table.floatright { margin: 0px 0px 0.5em 0.5em; border: 0px currentColor; } div.floatright p { font-style: italic; } div.floatleft { margin: 0px 0.5em 0.5em 0px; border: 0px currentColor; } table.floatleft { margin: 0px 0.5em 0.5em 0px; border: 0px currentColor; } div.floatleft p { font-style: italic; } div.thumb { width: auto; margin-bottom: 0.5em; background-color: transparent; } div.thumbinner { padding: 3px !important; border: 1px solid rgb(204, 204, 204); = text-align: center; overflow: hidden; font-size: 94%; background-color: = rgb(249, 249, 249); } html .thumbimage { border: 1px solid rgb(204, 204, 204); } html .thumbcaption { padding: 3px !important; border: currentColor; line-height: 1.4em; = font-size: 94%; } div.magnify { border: currentColor !important; } div.magnify a { border: currentColor !important; display: block; } div.magnify img { border: currentColor !important; display: block; } div.tright { margin: 0.5em 0px 1.3em 1.4em; } div.tleft { margin: 0.5em 1.4em 1.3em 0px; } img.thumbborder { border: 1px solid rgb(221, 221, 221); } #userlogin { margin: 0.5em; padding: 1.2em; border: 1px solid rgb(204, 204, 204); = float: left; } #userloginForm { margin: 0.5em; padding: 1.2em; border: 1px solid rgb(204, 204, 204); = float: left; } .catlinks { padding: 5px; border: 1px solid rgb(170, 170, 170); clear: both; = margin-top: 1em; background-color: rgb(249, 249, 249); } .usermessage { margin: 2em 0px 1em; padding: 0.5em 1em; border: 1px solid rgb(255, = 165, 0); color: black; font-weight: bold; vertical-align: middle; = background-color: rgb(255, 206, 123); } #siteNotice { margin: 0px; text-align: center; position: relative; } #localNotice { margin-bottom: 0.9em; } .firstHeading { line-height: 1.2em; padding-bottom: 0px; margin-bottom: 0.1em; } #firstHeading { line-height: 1.2em; padding-bottom: 0px; margin-bottom: 0.1em; } #siteSub { display: none; } #jump-to-nav { margin-top: -1.4em; margin-bottom: 1.4em; } #contentSub { margin: 0px 0px 1.4em 1em; width: auto; color: rgb(125, 125, 125); = line-height: 1.2em; font-size: 84%; } #contentSub2 { margin: 0px 0px 1.4em 1em; width: auto; color: rgb(125, 125, 125); = line-height: 1.2em; font-size: 84%; } span.subpages { display: block; } html { margin: 0px; padding: 0px; height: 100%; font-family: sans-serif; = font-size: 1em; } body { margin: 0px; padding: 0px; height: 100%; font-family: sans-serif; = font-size: 1em; } body { background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAABCAAAAAA6fptVAA= AACklEQVQIHWP4BgAA+AD3KTtPjAAAAABJRU5ErkJggg=3D=3D"); background-color: = rgb(243, 243, 243); } div#content { padding: 1em; color: black; margin-left: 10em; direction: ltr; = background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAABCAIAAACQd1PeAA= AADElEQVQIHWNYfv0nAASgAnhGoPmuAAAAAElFTkSuQmCC"); background-repeat: = repeat-y; background-color: white; } #mw-page-base { background-position: left bottom; height: 5em; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAsCAIAAAArRUU2AA= AAOklEQVQIHQXBAQEAAAjDIFz/zhdsCyE0y4SQya2RCSGWaYQ2Hc3ahFiINTIh1ly3NRohEwu= h0VmmER5oBCM5OFSTKwAAAABJRU5ErkJggg=3D=3D"); background-repeat: = repeat-x; background-color: white; } #mw-head-base { background-position: left bottom; height: 5em; margin-top: -5em; = margin-left: 10em; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAABCAIAAACQd1PeAA= AADElEQVQIHWNYfv0nAASgAnhGoPmuAAAAAElFTkSuQmCC"); background-repeat: = repeat-x; } div#mw-head { top: 0px; width: 100%; right: 0px; position: absolute; } div#mw-head h5 { margin: 0px; padding: 0px; } div.emptyPortlet { display: none; } #p-personal { top: 0.33em; right: 0.75em; position: absolute; } #p-personal h5 { display: none; } #p-personal ul { list-style: none; margin: 0px; padding-left: 10em; } #p-personal li { line-height: 1.12em; float: left; } #p-personal li { font-size: 0.75em; margin-top: 0.5em; margin-left: 0.75em; white-space: = nowrap; } #left-navigation { left: 10em; top: 2.5em; position: absolute; } #right-navigation { margin-top: 2.5em; float: right; } div.vectorTabs h5 { display: none; } div.vectorMenu h5 span { display: none; } div.vectorTabs { height: 2.5em; float: left; } div.vectorTabs { background-position: left bottom; padding-left: 1px; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAuCAIAAABmjeQ9AA= AARElEQVR4XmVO0QqAQAwy//+P87QjGUf0IDo3ZZREAD84eXkZ5YDBRQcFwpXv3kW1N4/GmZX= e326/xnf9ya8zzy/NtusBX91B1f55qzcAAAAASUVORK5CYII=3D"); = background-repeat: no-repeat; } div.vectorTabs ul { float: left; } div.vectorTabs ul { list-style: none; margin: 0px; padding: 0px; height: 100%; } div.vectorTabs ul li { float: left; } div.vectorTabs ul li { background-position: left bottom; margin: 0px; padding: 0px; height: = 100%; line-height: 1.12em; display: inline-block; white-space: nowrap; = background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAABkCAIAAADITs03AA= AAPElEQVR4XuWK2xEAIAzCsPvv5DSu0ahX3yv4wQVyGGCSvg2dnJvduT8sz/IwKOIfjCi2z/7= 6FhHeJcslVZQFLUC06LZ/AAAAAElFTkSuQmCC"); background-repeat: repeat-x; = background-color: rgb(243, 243, 243); } div.vectorTabs ul > li { display: block; } div.vectorTabs li.selected { background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAABkAQAAAABvV2fNAA= AADUlEQVQIHWNoYBgWEACJ5TIB0K9KcAAAAABJRU5ErkJggg=3D=3D"); } div.vectorTabs li a { height: 1.9em; color: rgb(6, 69, 173); padding-right: 0.5em; = padding-left: 0.5em; font-size: 0.8em; display: inline-block; cursor: = pointer; } div.vectorTabs li > a { display: block; } div.vectorTabs li.icon a { background-position: right bottom; background-repeat: no-repeat; } div.vectorTabs span a { padding-top: 1.25em; display: inline-block; } div.vectorTabs span > a { float: left; display: block; } div.vectorTabs span { background-position: right bottom; display: inline-block; = background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAuCAIAAABmjeQ9AA= AARElEQVR4XmVO0QqAQAwy//+P87QjGUf0IDo3ZZREAD84eXkZ5YDBRQcFwpXv3kW1N4/GmZX= e326/xnf9ya8zzy/NtusBX91B1f55qzcAAAAASUVORK5CYII=3D"); = background-repeat: no-repeat; } div.vectorTabs li.selected a { color: rgb(51, 51, 51); text-decoration: none; } div.vectorTabs li.selected a:visited { color: rgb(51, 51, 51); text-decoration: none; } div.vectorTabs li.new a { color: rgb(165, 88, 88); } div.vectorTabs li.new a:visited { color: rgb(165, 88, 88); } div.vectorMenu { background-position: 100% 60%; float: left; cursor: pointer; direction: = ltr; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAABYAAAAQBAMAAADgw5IVAA= AAD1BMVEXz+fuamprd3d15eXmxsbG+1Cf6AAAAAXRSTlMAQObYZgAAACZJREFUCFtjYKAaYBI= 2NjYWgnIUjY0NFRASMGGgBFwYKOEEZ5ILAABbAqkPJqHJAAAAAElFTkSuQmCC"); = background-repeat: no-repeat; } div.vectorMenuFocus { background-position: 100% 60%; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAABYAAAAQBAMAAADgw5IVAA= AAD1BMVEWampqSkpKtra2/v7/j4+Mm0C2mAAAAAXRSTlMAQObYZgAAACZJREFUCFtjYKAaYFE= UFBRUgXKcBAWFHBASMGGgBFwYKGECZ5ILANu8AllBO68YAAAAAElFTkSuQmCC"); } body.rtl div.vectorMenu { direction: rtl; } div#mw-head div.vectorMenu h5 { float: left; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAuCAIAAABmjeQ9AA= AARElEQVR4XmVO0QqAQAwy//+P87QjGUf0IDo3ZZREAD84eXkZ5YDBRQcFwpXv3kW1N4/GmZX= e326/xnf9ya8zzy/NtusBX91B1f55qzcAAAAASUVORK5CYII=3D"); = background-repeat: no-repeat; } div#mw-head div.vectorMenu h5 { background-position: left bottom; margin-left: -1px; } div#mw-head div.vectorMenu > h5 { background-image: none; } div#mw-head div.vectorMenu h4 { border: currentColor; padding-top: 1.37em; padding-left: 0.5em; = font-size: 0.8em; font-weight: normal; float: left; display: = inline-block; } div.vectorMenu h5 a { width: 24px; height: 2.5em; text-decoration: none; display: = inline-block; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAuCAIAAABmjeQ9AA= AARElEQVR4XmVO0QqAQAwy//+P87QjGUf0IDo3ZZREAD84eXkZ5YDBRQcFwpXv3kW1N4/GmZX= e326/xnf9ya8zzy/NtusBX91B1f55qzcAAAAASUVORK5CYII=3D"); = background-repeat: no-repeat; } div.vectorMenu h5 a { background-position: right bottom; } div.vectorMenu h5 > a { display: block; } div.vectorMenu div.menu { text-align: left; clear: both; display: none; position: relative; } body.rtl div.vectorMenu div.menu { margin-left: 24px; } body.rtl div.vectorMenu > div.menu { margin-left: auto; } div.vectorMenu:hover div.menu { display: block; } div.vectorMenu div.menuForceShow { display: block; } div.vectorMenu ul { border-width: 0px 1px 1px; border-style: solid; border-color: silver; = list-style: none; margin: 0px 0px 0px -1px; padding: 0px; text-align: = left; position: absolute; background-color: white; } div.vectorMenu li { margin: 0px; padding: 0px; text-align: left; line-height: 1em; } div.vectorMenu li a { padding: 0.5em; color: rgb(6, 69, 173); font-size: 0.8em; display: = inline-block; white-space: nowrap; cursor: pointer; } div.vectorMenu li > a { display: block; } div.vectorMenu li.selected a { color: rgb(51, 51, 51); text-decoration: none; } div.vectorMenu li.selected a:visited { color: rgb(51, 51, 51); text-decoration: none; } #p-search h5 { display: none; } #p-search { float: left; } #p-search { margin-right: 0.5em; margin-left: 0.5em; } #p-search form { margin: 0.4em 0px 0px; } #p-search input { margin: 0.4em 0px 0px; } div#simpleSearch { border: 1px solid rgb(170, 170, 170); width: 14em; height: 1.4em; = color: black; margin-top: 0.65em; display: block; position: relative; = min-height: 1px; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAQCAIAAABY/YLgAA= AAJUlEQVQIHQXBsQEAAAjDoND/73UWdnerhmHVsDQZJrNWVg3Dqge6bgMe6bejNAAAAABJRU5= ErkJggg=3D=3D"); background-repeat: repeat-x; background-color: white; } div#simpleSearch label { top: 0.25em; font-size: 13px; direction: ltr; } div#simpleSearch input { color: black; direction: ltr; } div#simpleSearch input:focus { =09 } div#simpleSearch input.placeholder { color: rgb(153, 153, 153); } div#simpleSearch input#searchInput { margin: 0px; padding: 0.2em 0px 0.2em 0.2em; border: currentColor; = left: 0px; top: 0px; width: 90%; font-size: 13px; position: absolute; = direction: ltr; background-color: transparent; } div#simpleSearch button#searchButton { margin: 0px; padding: 0.3em 0.4em 0.2em 0px; border: currentColor; top: = 0px; width: 10%; right: 0px; position: absolute; cursor: pointer; = background-image: none; background-color: transparent; } div#simpleSearch button#searchButton img { margin: -3px 0px 0px; padding: 0px; border: currentColor; } div#simpleSearch button#searchButton > img { margin: 0px; } div#mw-panel { left: 0px; top: 160px; width: 10em; padding-top: 1em; position: = absolute; } div#mw-panel div.portal { padding-bottom: 1.5em; direction: ltr; } div#mw-panel div.portal h5 { padding: 0px 0.25em 0.25em 1.75em; border: currentColor; color: rgb(68, = 68, 68); font-size: 0.75em; font-weight: normal; cursor: default; } div#mw-panel div.portal div.body { margin: 0px 0px 0px 1.25em; padding-top: 0.5em; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAIwAAAABCAAAAAAphRnkAA= AAJ0lEQVQIW7XFsQEAIAyAMPD/b7uLWz8wS5youFW1UREfiIpH1Q2VBz7fGPS1dOGeAAAAAEl= FTkSuQmCC"); background-repeat: no-repeat; } div#mw-panel div.portal div.body ul { list-style: none; margin: 0px; padding: 0px; } div#mw-panel div.portal div.body ul li { margin: 0px; padding: 0px 0px 0.5em; line-height: 1.12em; overflow: = hidden; font-size: 0.75em; } div#mw-panel div.portal div.body ul li a { color: rgb(6, 69, 173); } div#mw-panel div.portal div.body ul li a:visited { color: rgb(11, 0, 128); } div#footer { padding: 0.75em; margin-top: 0px; margin-left: 10em; direction: ltr; = background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAABCAIAAACQd1PeAA= AADElEQVQIHWNYfv0nAASgAnhGoPmuAAAAAElFTkSuQmCC"); background-repeat: = repeat-x; } div#footer ul { list-style: none; margin: 0px; padding: 0px; } div#footer ul li { margin: 0px; padding: 0.5em 0px; color: rgb(51, 51, 51); font-size: = 0.7em; } div#footer #footer-icons { float: right; } body.ltr div#footer #footer-places { float: left; } div#footer #footer-info li { line-height: 1.4em; } div#footer #footer-icons li { text-align: right; line-height: 2em; margin-left: 0.5em; float: left; } div#footer #footer-places li { line-height: 2em; margin-right: 1em; float: left; } #p-logo { left: 0px; top: -160px; width: 10em; height: 160px; position: absolute; } #p-logo a { background-position: center; width: 10em; height: 160px; = text-decoration: none; display: block; background-repeat: no-repeat; } #preftoc { background-position: left bottom; margin: 0px !important; padding: 0px = !important; width: 100%; clear: both; float: left; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAhCAMAAAAvZzGNAA= AAYFBMVEUAAAD+/v79/f38/Pzt7e36+vr4+Pj39/f19fXz8/Py8vLw8PD7+/vq6uro6Ojm5ub= l5eXj4+Ph4eHf39/d3d3b29vZ2dnY2NjW1tbU1NTT09PR0dHQ0NDPz8/Ozs7Nzc0koCh/AAAA= AXRSTlMAQObYZgAAABtJREFUCFtjYmBiBENOpp9QFgT+ZGIAijEShgBszwJih9jAuAAAAABJR= U5ErkJggg=3D=3D"); background-repeat: no-repeat; } #preftoc li { background-position: right bottom; margin: 0px; padding: 0px 1px 0px = 0px; height: 2.25em; float: left; list-style-type: none; = list-style-image: none; white-space: nowrap; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAhCAMAAAAvZzGNAA= AAYFBMVEUAAAD+/v79/f38/Pzt7e36+vr4+Pj39/f19fXz8/Py8vLw8PD7+/vq6uro6Ojm5ub= l5eXj4+Ph4eHf39/d3d3b29vZ2dnY2NjW1tbU1NTT09PR0dHQ0NDPz8/Ozs7Nzc0koCh/AAAA= AXRSTlMAQObYZgAAABtJREFUCFtjYmBiBENOpp9QFgT+ZGIAijEShgBszwJih9jAuAAAAABJR= U5ErkJggg=3D=3D"); background-repeat: no-repeat; } #preftoc li:first-child { margin-left: 1px; } #preftoc a { padding: 0.5em; color: rgb(6, 69, 173); font-size: 0.9em; = text-decoration: none; display: inline-block; position: relative; = background-image: none; } #preftoc a:active { padding: 0.5em; color: rgb(6, 69, 173); font-size: 0.9em; = text-decoration: none; display: inline-block; position: relative; = background-image: none; } #preftoc a:hover { text-decoration: underline; } #preftoc a:focus { text-decoration: underline; } #preftoc li.selected a { background-position: bottom; color: rgb(51, 51, 51); text-decoration: = none; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAAhCAQAAACysAk0AA= AAHElEQVQIW2P6z8DE8J+JgQGKQFwMhCaIn4shCAC27Qc86OAoxwAAAABJRU5ErkJggg=3D=3D= "); background-repeat: repeat-x; } #preferences { margin: -2px 0px 0px; border: 1px solid rgb(204, 204, 204); width: = 100%; clear: both; float: left; background-image: = url("data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAAEAAAABCAAAAAA6fptVAA= AACklEQVQIHWP4BQAA/AD7oRDZ1QAAAABJRU5ErkJggg=3D=3D"); background-color: = rgb(249, 249, 249); } #preferences fieldset { border-width: 1px medium medium; border-style: solid none none; = border-color: rgb(204, 204, 204) currentColor currentColor; } #preferences fieldset.prefsection { margin: 1em; padding: 0px; border: currentColor; } #preferences legend { color: rgb(102, 102, 102); } #preferences fieldset.prefsection legend.mainLegend { display: 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